作　　者： ; ;

机构地区： 山东大学生命科学学院生物科学系

出　　处： 《山东大学学报（自然科学版）》 1992年第2期207-214,共8页

摘　　要： 野生大豆绒毡层属分泌型,在发育过程中始终是单核。绒毡层细胞是花粉外壁孢粉素的主要来源;在不同的发育时期,绒毡层细胞以不同的方式将孢粉素传递到花粉囊中:(1)减数分裂时期,绒毡层细胞的内质网形成大量膜包原生质小泡,这可能是绒毡层细胞适应孢粉素大量合成而形成的一种结构;此时只有少量乌氏体被排到径向质膜外方。(2)四分体时期,绒毡层细胞分泌出大量无定形的孢粉素团块进入花粉囊中,而成为花粉外壁孢粉素的最主要来源。(3)单核花粉时期,绒毡层细胞内切向质膜形成港湾结构,原乌氏体在其中发育成乌氏体;这些后期形成的乌氏体可能与花粉外壁覆盖层的进一步发育有关。 The development of the tapetum in Glycine soja has beer.examined atthe ultrastructural level.The grandular type tapetum remain uninucleatethroughout its development.The sporopollenin of pollen exine is mainly comefrom the tapetal cell,but it seem to have different ways for sporopollenintransporting during the development of microspore.(1).Mciotic stage:A greatdeal of membrane-bound protoplasmic vesicles,which are formed by RER,appear in the cytoplasm of tapetal cell to fit for synthesizing sporopollenin.The pro-Ubisch bodies are found in the vesicles and later they are extrudedto the outside of radial plasmolemma of tapetal cell.(2).Tetrad stage:Tapetal cell secrete a great deal of medium electron dense,shapeless materialwhich is probably as the mainly source of sporopollenin,of pollen exine.(3).Uninucleate pollen stage,The inner tangential plasmolemma of tapetalcell form lot of embayment-like structure in which the pro-Ubisch bodiescan be found and with continued sporopollenin depositing,they develop intoUbisch bodies and leave the embayment-like structure to the tapetal membr-ance.Following the breakdown of tapetal cell,they enter the locule and maypresent another way of sporopollenin tr(?)

关 键 词： 野生大豆 绒毡层细胞 超微结构

领　　域： [生物学]